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为解决视觉皮层中物体类别选择性区域的组织原则问题，研究人员开展了一项关于动作维度如何塑造视觉皮层地形图的研究。他们通过fMRI实验发现，左侧外侧枕颞皮层(LOTC)存在一个基于动作属性的地形梯度，而腹侧枕颞皮层(VOTC)则主要由生命性(animacy)主导。